
NOTES FROM NATURE
By Jerry Toll
The Breeding Season
The first warm days of spring are finally upon us.
Robins, Red-winged Blackbirds, and waterfowl have returned in profusion.
Come June, most of the birds in our area will be nesting. Not all species
follow this pattern, though. The owls will have completed their breeding
cycle by the time the neotropical birds begin theirs. There are overriding
concepts that clarify why different species time their breeding cycles in
different ways.
Energy Demands
From an ecological point of view, on both the individual
and species level, energy demands largely determine the ability to survive
and reproduce. The ability to obtain food, retain the energy, and pass
energy on to the next generation is the foundation of survival.
The following quoted passages are taken from The
Birder's Handbook by Erlich, Dobkin and Wheye.
"Evolution generally adjusts the timing of avian breeding
seasons to produce the most young. Nebraska is in the temperate region.
From this region to the subarctic and arctic regions, the main factor is
availability of food.
Abundant nourishment is needed, not only by nestlings and
juveniles, but by the increased energy demands on breeding adults. For
females it's the energetic demand of producing eggs. For males it's the
increased energy demands of vigorous displays and defending territories.
One or both adults participate in building a nest, foraging for more than
one individual (mate and/or chicks), and in some cases territorial defense
or guarding young from predators. "sense that it is time to start breeding by the
lengthening of the days as spring approaches."
For the neotropical species who spend the majority of
their lives in the tropics, migration is a necessary part of the breeding
cycle. For some it's the lengthening days; for others it's the internal
calendars that tell them conditions will be good for breeding when they
arrive.
Even many tropical birds time breeding to environmental
changes. Some may time breeding to the prevalent wet or dry seasons.
Others may breed irrespective of environmental changes because some
influences like food supply are less limiting in the tropics.
"Evolution seems to have latched on to the length of
daylight as a timer of activities, since it is a signal that can be used to forecast future events." For instance, the
Snow Geese who migrated through Omaha on March 14 could have waited
another couple of weeks until the lakes opened up and the snow melted.
This would have made their migration easier by making available more food
and safe roosting, but it wouldn't have allowed them enough time on the
breeding ground to maximize breeding success. The length of daylight told
them it was necessary to press on, irrespective of the hardship.
Factors like food supply and weather may fine tune day
length. If Red-winged Blackbirds are experimentally provided with abundant
food, they will begin laying eggs three weeks earlier than birds without
an abundant supply.
Some species become so specialized in their breeding
requirements that extinction looms near. Others, though more general in
breeding strategy, are threatened by the rapid changes brought on by human
influences on the environment.
This article describes broad concepts about ways in which
birds time breeding. At every level, birds have found unique and subtle
strategies to accomplish successful breeding.

Previous Notes from Nature:
01/24/08